Publications (prior to 2015)

Gilbert, B., Tunney, T.D., McCann, K.S., DeLong, J.P., Vasseur, D.A., Savage, V., Shurin, J.B., Dell, A.I., Barton, B.T., Harley, C.D.G., Kharouba^G, H.M., Kratina, P., Blanchard, J.L., Clements, C., Winder, M., Greig, H.S., & O’Connor, M.I. 2014. A bioenergetic framework for the temperature dependence of trophic interactions. Ecology Letters 17:902-914.

Helmuth, B., Mieszkowska, N., Connell, S., Dong, Y., Harley, C.D.G., Lima, F., Sara, G., Williams, G., & Russell, B. 2014. Beyond long-term averages: making biological sense of a rapidly changing world. Climate Change Responses 1:10-20.

Tallis, H., Lubchenco, J., & 238 co-signatories. 2014. Working together: A call for inclusive conservation. Nature 515:27-28. [Comment in Nature.]

Vasseur, D.A., DeLong, J.P., Gilbert, B., Grieg, H., Harley, C.D.G., McCann, K.S., O’Connor, M.I., Tunney, T., & Savage, V. 2014. Temperature variation puts species at greater risk under global change. Proceedings of the Royal Society B 281:20132612.

Demes^G, K.W., Harley, C.D.G., Anderson, L.M., & Carrington, E. 2013. Shifts in morphological and mechanical traits compensate for performance costs of reproduction in a wave-swept seaweed. Journal of Ecology 101:963-970.

 Demes^G, K.W., Pruitt, J.N., Harley, C.D.G., & Carrington, E. 2013. Survival of the weakest: Increased frond mechanical strength in a wave-swept kelp inhibits self-pruning and increases mortality. Functional Ecology 27:439-445. [Winner of the 2013 Haldane Prize for best paper by a young author published in Functional Ecology.]

Harley, C.D.G. 2013. Linking ecomechanics and ecophysiology to interspecific interactions and community dynamics. Annals of the New York Academy of Sciences 1297:73-82.

Harley, C.D.G., Anderson^G, K.M., Lebreton^U, C.A.-M., MacKay^U, A., Ayala-Díaz, M., Chong^U, S.L., Pond^U, L.M., Amerongen Maddison^U, J.H., Hung^U, B.H.C., Iverson^U, S.L., & Wong^U, D.C.M. 2013. The introduction of Littorina littorea to British Columbia, Canada: potential impacts and the importance of biotic resistance. Marine Biology 160:1529-1541. [feature article]

Singh^G, G., Markel, R., Martone, R., Salomon, A., Harley, C.D.G., and Chan, K. 2013. Sea otters homogenize mussel beds and reduce habitat provisioning in a rocky intertidal ecosystem. PLOS ONE 8(5):e65435.

Harley, C.D.G., Anderson^G, K.M., Demes^G, K.W., Jorve^G, J.P., Kordas^G, R.L., Coyle^U, T.A., & Graham, M.H. 2012. Effects of climate change on global seaweed communities. Journal of Phycology 48:1064-1078. [journal cover photo; most highly cited paper (of 294) published in this journal in 2012]

Russell, B.D., Harley, C.D.G., Wernberg, T., Mieszkowska, N., Widdicombe, S., Hall-Spencer, J.M., & Connell, S.D. 2012. Predicting ecosystem shifts requires new approaches that integrate the effects of climate change across entire systems. Biology Letters 8:164-166.

Crim^G, R.N., Sunday, J.M., & Harley, C.D.G. 2011. Elevated seawater CO₂ concentrations impair larval development and reduce larval survival in endangered northern abalone (Haliotis kamtschatkana). Journal of Experimental Marine Biology and Ecology 400:272-277.

Harley, C.D.G. 2011. Climate change, keystone predation, and biodiversity loss. Science 334:1124-1127. [Twice recommended by Faculty of 1000]

Harley, C.D.G., & O’Riley^U, J. 2011. Non-linear density dependent effects of an intertidal ecosystem engineer. Oecologia 166:531-541.

Kordas^G, R.L., Harley, C.D.G., & O’Connor, M.I. 2011. Community ecology in a warming world: the influence of temperature on interspecific interactions in marine systems. Journal of Experimental Marine Biology and Ecology 400:218-226. [Top 5 in citations of >2500 papers published in this journal in the last 10 years.]

Reuter, K.E., Lotterhos, K.E., Crim^G, R.N., Thompson^U, C.A., & Harley, C.D.G. 2011. Ocean acidification increases sperm limitation and risk of polyspermy in the red sea urchin Strongylocentrotus franciscanus. Global Change Biology 17:163-171.

Sunday, J.M., Crim^G, R., Harley, C.D.G., & Hart, M.W. 2011. Quantifying rates of evolutionary adaptation in response to ocean acidification. PLOS ONE 6(8):e22881. doi:10.1371/journal.pone.0022881. [Ranks in top 1% in citations for this journal.]

 Bates^P, A., Stickle, W., & Harley, C.D.G. 2010. Impact of temperature on the parasitic association between a sperm-feeding scuticociliate and Pacific west coast sea stars. Journal of Experimental Marine Biology and Ecology 384:44-50.

Nienhuis^G, S., Palmer, A.R., & Harley, C.D.G. 2010. Elevated CO₂ affects shell dissolution rate but not calcification rate in a marine snail. Proceedings of the Royal Society B 277:2553-2558.

Bates^P, A., Hilton^U, B.J., & Harley, C.D.G. 2009. Effects of seasonal variation and temperature on wasting disease in the keystone predatory sea star, Pisaster ochraceus. Diseases of Aquatic Organisms 86:245-251.

Held^U, M., & Harley, C.D.G. 2009. Responses to low salinity by the sea star Pisaster ochraceus from high- and low-salinity populations. Invertebrate Biology 128:381-390.

Denny, M.W., Hunt, L.J.H., Miller, L.P., & Harley, C.D.G. 2009. On the prediction of extreme ecological events. Ecological Monographs 79:397-421.

Gooding^G, R.A., Harley, C.D.G., & Tang^U, E. 2009. Elevated water temperature and carbon dioxide concentration increase the growth of a keystone echinoderm. Proceedings of the National Academy of Sciences USA 106:9316-9321.

Harley, C.D.G., Denny, M.W., Mach, K., & Miller, L.P. 2009. Thermal stress and morphological adaptations in limpets. Functional Ecology 23:292-301.

Harley, C.D.G., & Paine, R.T. 2009. Contingencies and compounded rare perturbations dictate sudden distributional shifts during periods of gradual climate change. Proceedings of the National Academy of Sciences USA 106:11172-11176.

Miller, L.P., Harley, C.D.G., & Denny, M.W. 2009. The role of temperature and desiccation stress in limiting the local-scale distribution of the owl limpet, Lottia gigantea. Functional Ecology 23:756-767.

Harley, C.D.G. 2008. Tidal dynamics, topography, and temperature-mediated mass mortalities on rocky shores. Marine Ecology Progress Series 371:37-46.

Morelissen^G, B., & Harley, C.D.G. 2007.The effects of thermal stress on producers, consumers, and plant-herbivore interactions in an intertidal community. Journal of Experimental Marine Biology and Ecology 348:162-173.

Denny, M.W., & Harley, C.D.G. 2006. Hot limpets: predicting body temperature in a conductance-mediated thermal system. Journal of Experimental Biology 209:2409-2419.

Denny, M.W., Miller, L.P., & Harley, C.D.G. 2006. Thermal stress on intertidal limpets: long-term hindcasts and lethal limits. Journal of Experimental Biology 209:2420-2431.

Gilman, S.E., Harley, C.D.G., Strickland, D.C., Vanderstraeten, O., O’Donnell, M.J., & Helmuth, B. 2006. Evaluation of effective shore level as a method of characterizing intertidal wave exposure regimes. Limnology & Oceanography Methods 4:448-457.

Harley, C.D.G. 2006. Direct and indirect effects of ecosystem engineering and herbivory on intertidal community structure. Marine Ecology Progress Series 317:29-39.

Harley, C.D.G., Hughes, A.R., Hultgren, K.M., Miner, B.G., Sorte, C.J.B., Thornber, C.S., Rodriguez, L.F., Tomanek, L., & Williams, S. L. 2006. The impact of climate change in coastal marine systems. Ecology Letters 9:228-241. [journal cover photo, 2^nd most highly cited paper (of 150) published in Ecology Letters in 2006, cited >1000 times]

Harley, C.D.G., Pankey, M.S., Wares, J.P., Wonham, M.J., & Grosberg, R. 2006. Color polymorphism and genetic structure in the sea star Pisaster ochraceus. Biological Bulletin 211:248-262. [journal cover photo]

Helmuth, B., Broitman, B., Blanchette, C.A., Gilman, S., Halpin, P., Harley, C.D.G., O’Donnell, M., Hofmann, G.E., Menge, B., & Strickland, D. 2006. Mosaic patterns of thermal stress in the rocky intertidal zone: implications for climate change. Ecological Monographs 76:461-479. [journal cover photo]

Wonham, M.J., O’Connor^U, M., & Harley, C.D.G. 2005. Enhancement of introduced and native species by a dominant invader. Marine Ecology Progress Series 289:109-116.

Harley, C.D.G. & Rogers-Bennett, L. 2004. The potential synergistic effects of climate change and fishing pressure on exploited invertebrates on rocky intertidal shores. California Cooperative Oceanic Fisheries Investigations Reports 45:98-110.

 Denny, M.W., Helmuth, B., Leonard, G.H., Harley, C.D.G., Hunt, L.J.H., & Nelson, E.K. 2004. Quantifying scale in ecology: lessons from a wave-swept shore. Ecological Monographs 74:513-532. [journal cover photo]